Virtually all game companies use Trunk Based Development, in my experience, including many very large studios. (Outside of game engine development, which isn't really done by game studios anymore with a few exceptions)
I suspect you'll find that two of those numbers are decided by committee, one by a human, and the other by the build system.
There's good odds that it's the 26836th build, the 4045th snapshot, and the second time they've broken the file format. (or the 2nd expansion, or both)